Plus500 Brumbies prop partners, Scott Sio and Allan Alaalatoa will once again fill the bookend jerseys for the Wallabies in their crunch pool D clash against Wales.
The powerful pair will look to lock down a Welsh pack that can be very potent on their day, fellow Brumby James Slipper will also have an opportunity to contribute as a finisher.
Rory Arnold retains his place in the second row while David Pocock will wear six for the second game in a row.
Elsewhere, former Brumbies star Adam Ashley-Cooper starts on the wing in place of suspended Reece Hodge, becoming of just a handful of players who have performed at four Rugby World Cups.
Australia last met Wales on the 2018 Spring Tour in Cardiff when the hosts dug deep for a gritty 9-6 win at Millennium Stadium.
The Wallabies and Wales have enjoyed some memorable battles at the Rugby World Cup, having met six times at Rugby’s showpiece (Australia W5, L1).
The Wallabies will next take on Uruguay in Oita on October 5 before shifting north again to Shizuoka for the final Pool D clash against Georgia the following weekend. The two highest-ranked teams at the end of the pool stage will advance to the Quarter Finals.
Foxtel will showcase every Rugby World Cup game LIVE across two dedicated channels with no ad-breaks during play and in 4K Ultra High Definition. All games will also be streamed LIVE and On Demand on Kayo Sports.
Ten of the biggest matches from the 2019 Rugby World Cup will air LIVE and free on Network 10, WIN Network and tenplay.com.au, including every Australia match right up to and including the Final.
2019 Rugby World Cup
Wallabies 39 v Fiji 21 at Sapporo Dome, Sapporo, Japan, Saturday 21 September at 2.45pm AEST
Wallabies v Wales at Ajinomoto Stadium, Tokyo, Sunday 29 September at 5.45pm AEST
Wallabies v Uruguay at Oita Bank Dome, Oita, Saturday 5 October at 3.15pm AEST
Wallabies v Georgia at Shizuoka Stadium, Shizuoka, Friday 11 October at 9.15pm AEST